This particular PDF no longer prints in Karmic

[Matthew Paul Thomas]

Evince 2.28.1, Ubuntu Karmic; Xerox WorkCentre 7228

The attached PDF prints fine from Evince in Ubuntu 9.04. In Ubuntu Karmic, however, once it is sent to the printer the printer beeps once and does nothing else.

The same problem happens with Evince on two separate computers.
Evince successfully prints other PDFs on the same computers.
The same document prints successfully using Adobe Reader 9 from the same computer.

ProblemType: Bug
Architecture: i386
CupsErrorLog: E [20/Oct/2009:18:01:32 +0100] Unable to remove temporary file "/var/spool/cups/tmp/.hplip" - Is a directory
Date: Tue Oct 20 18:20:43 2009
DistroRelease: Ubuntu 9.10
MachineType: Apple Inc. MacBook4,1
NonfreeKernelModules: wl
Package: cups 1.4.1-5ubuntu2
Papersize: a4
PpdFiles:
 HP-Color-LaserJet-3600: HP Color LaserJet 3600 Foomatic/pxljr
 Deskjet-F300-series: HP Deskjet f300 Series hpijs, 3.9.8
ProcCmdLine: BOOT_IMAGE=/boot/vmlinuz-2.6.31-14-generic root=UUID=11cc112e-2fe8-415f-91d5-fd68b0f8dc1f ro quiet splash
ProcEnviron:
 LANGUAGE=en_NZ.UTF-8
 PATH=(custom, user)
 LANG=en_NZ.UTF-8
 SHELL=/bin/bash
ProcVersionSignature: Ubuntu 2.6.31-14.47-generic
SourcePackage: cups
Uname: Linux 2.6.31-14-generic i686
dmi.bios.date: 02/09/08
dmi.bios.vendor: Apple Inc.
dmi.bios.version: MB41.88Z.00C1.B00.0802091535
dmi.board.asset.tag: Base Board Asset Tag
dmi.board.name: Mac-F22788A9
dmi.board.vendor: Apple Inc.
dmi.board.version: PVT
dmi.chassis.asset.tag: Asset Tag#
dmi.chassis.type: 2
dmi.chassis.vendor: Apple Inc.
dmi.chassis.version: Mac-F22788A9
dmi.modalias: dmi:bvnAppleInc.:bvrMB41.88Z.00C1.B00.0802091535:bd02/09/08:svnAppleInc.:pnMacBook4,1:pvr1.0:rvnAppleInc.:rnMac-F22788A9:rvrPVT:cvnAppleInc.:ct2:cvrMac-F22788A9:
dmi.product.name: MacBook4,1
dmi.product.version: 1.0
dmi.sys.vendor: Apple Inc.